Natufians and Neolithic Levantines lack African admixture?

Whilst Neolithic Levantines and Natufians seem to have a good amount of what looks to be Y-DNA E-M35 (specifically 2 E-Z830 samples among the Natufians) and seemingly no Y-DNA J so far- :

Natufians (Epipaleolithic Levant):

I0861: E1b1b1b2(x E1b1b1b2a, E1b1b1b2b)
I1069: E1b1(xE1b1a1, E1b1b1b1)
I1072: E1b1b1b2(xE1b1b1b2a, E1b1b1b2b)
I1685: CT
I1690: CT

Levant_Neolithic:

I0867: H2 (PPNB)
I1414: E(xE2, E1a, E1b1a1a1c2c3b1, E1b1b1b1a1, E1b1b1b2b) (PPNB)
I1415: E1b1b1 (PPNB)
I1416: CT (PPNB)
I1707: T(xT1a1, T1a2a) (PPNB)
I1710: E1b1b1(x E1b1b1b1a1, E1b1b1a1b1, E1b1b1a1b2, E1b1b1b2a1c) (PPNB)
I1727: CT(xE, G, J, LT, R, Q1a, Q1b) (PPNB)
I1700: CT (PPNC)


-they don't seem to show any affinities for African populations (ones who lack West Eurasian admixture or have negligible amounts of such admixture) on an autosomal level according to this Pre-print:

"However, no affinity of Natufians to sub-Saharan Africans is evident in our genome-wide analysis, as present-day sub-Saharan Africans do not share more alleles with Natufians than with other ancient Eurasians (Extended Data Table 1). (We could not test for a link to present-day North Africans, who owe most of their ancestry to back-migration from Eurasia)."

This is quite interesting, truth be told. It implies that, despite seeming clearly old, the African admixture in the Levant is perhaps, at least, Post-Neolithic? Some of it (particularly non-East African cluster-related admixture), as I've noted in the past, is definitely owed to the Arab Slave Trade but some of it does also seem rather ancient.  

As for PCA (Principal Component Analysis) positions; Natufians in particular look to cluster essentially to the direct east of Anatolian Neolithic and Early European Farmers (and somewhat southwards of some of them) whilst Levantine Neolithic Farmers seem less Basal Eurasian and more northern shifted than their Epipaleolithic predecessors. I'll be more interested in seeing David over at Eurogenes throw these samples into his own Pan-West Eurasia PCA and a global PCA once the genomes are made publicly available when this paper is published in a peer-reviewed journal (a month or less, not sure). The PCAs in these studies can be a bit wonky due to issues like projection-bias.

I'll be interested in seeing if David and other 3rd parties will either confirm or deny that Natufians lack African admixture (whatever Basal Eurasian turns out to be aside).

One somewhat off-topic matter to note is that it seems as though we've more or less discovered a close equivalent to David's "ENF"/Near Eastern cluster from his old Fateful Triangle~K=8 model, a cluster also noted by Lazaridis and company prior to the release of this pre-print. 

The Natufians and Neolithic Levantines are essentially dominated by the blue ADMIXTURE cluster that seems to make up most of the ancestry in the Neolithic Anatolians and Early European Farmers. This seems, to me, like we're seeing Southwest Asian/ENF/Near Eastern in the flesh. The way it's differentiated from Neolithic Iranians is intriguing though given that Natufians (Epipaleolithic Levantines) and Neolithic Iranians are comparable in terms of Basal Eurasian ancestry; it must be the non-Basal Eurasian ancestry that's causing the differentiation or there's just really significant genetic drift at play here or both.

In this model above, where Natufians and the Hotu cave Iranian were not included, the Caucasus Hunter-Gatherer (CHG) samples seem to carry notable EHG/ANE-related ancestry as prior analyses like David's K=8 ADMIXTURE run might've implied. Initially, I figured what was differentiating the Neolithic Iranians from Natufians was perhaps ANE-related admixture in the former but that might not really be the case.

As I said in a prior post; things will become more clear when third-parties like David over at Eurogenes get their hands on these genomes once the paper's published at a peer-reviewed journal and also once we have even more ancient genomes someday.

Reference List:

1. The genetic structure of the world's first farmers, Lazaridis et al. 2016 

Note:

1. I also must add that we should be cautious given that these are just 6 Natufian samples from one area. There might be some variation with future samples from other areas in regards to Y-DNA although I doubt there will be very much variation in terms of autosomal DNA given their similarity to Levantines from later periods.